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galactitol dulcitol sugar alcohol reduction galactose sweet d-galactitol galactokinase deficiency galactosemia cataracts aldose reductase galactose-1-phosphate uridylyltransferase autosomal recessive disorder galactose-1-phosphate metabolism rendering metabolite biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous contractor human metabolome database hmdb molecule molecular structural structure formula skeletal geometry ball stick