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diptera large order estimated 000 species including horseflies crane flies hoverflies mosquitoes one major insect orders considerable ecological human importance important pollinators second bees hymenopteran relatives evolutionarily earliest responsible early plant pollination fruit model organisms research abundant found terrestrial habitats world antarctica include familiar insects house blow gnats black midges formally actual diversity greater parts studied intensively greatest gall forming principally family cecidomyiidae close fly caught spider leaf