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poly adpribose polymerase parp1 dna damage detection protein target cancer drug development adp ribose parp repair programmed cell death caspase single strand break ssb par apoptosis lifespan longevity inhibitor tumor nick talazoparib olaparib rucaparib veliparib ribosyltransferase podb4dqy structure peptide amino acid molecule polypeptide polymer atoms science scientific disease health molecular model cartoon representation combined surfaces